A list of Philippine mosquitoes (Diptera: Culicidae).

Mosquito Systematics Vol. 17(1) 1985 1

A List of Philippine Mosquitoes
(Diptera: Culicidae)

1 2 1

Adela Cagampang-Ramos , Robert J. McKenna , and Dennis D. Pinkovsky

ABSTRACT. An updated list of the mosquitoes recorded from the Republic of
the Philippines is provided. The list contains 279 species, subspecies, and
varieties. Geographic distributions and pertinent references are incorporated
within the taxonomic listing.

The number of Philippine mosquitoes recorded in the literature has increased
from the two species noted by Osten-Sacken (1882) to the 279 species, subspecies,
and varieties listed here. The Philippine Culicidae were last reviewed by
Basio (1971, 1974). Basio's works are not well known because they were
published in journals with a limited distribution. Since Basio's publications,
there have been significant publications in culicid taxonomy which have affected
the faunal list of the Philippines: Abercrombie (1977); Harrison and Scanlon
(1974, 1975); Huang (1978b, 1979); Knight (1978); Knight and Stone (1977);
Mattingly (1980); Miyagi (1980); Miyagi , Toma and Tsukamoto (1983); Miyagi ,
Toma and Rivera (1983); Peyton (1977); Ramalingam (1974); Reinert (1970, 1974,
1976a, 1976b, 1981, 1982); and Sirivanakarn (1972, 1976, 1977a). Our list
includes these changes, reflecting 35 specific or subspecific additions of which
15 are new descriptions, 7 are new records, and 17 are resurrections from
synonymy. Supraspecific additions are the genus Mimomyia (Mattingly 1971) and
the subgenera Belkinius (Reinert 1982), Ingvamia (White 1974), Raohionotomyia
(Mattingly 1980), and Tricholeptomyia (Mattingly 1980). Forty species have
been reassigned to these new taxa. Thirteen species and the subgenera
Raohisouva, Ravenalities 3 and Skusea listed by Basio have been deleted. Pertinent
literature and Philippine distribution, based primarily on published reports and
locally examined specimens, are included within our list. New locations,
previously unpublished, are indicated by an asterisk. Other (worldwide)
distribution for the listed species is available in the Catalogue of the
Mosquitoes of the World (Knight and Stone 1977) and the supplement (Knight
1978) to that publication. Format for citing geographic distribution is as
follows: specific locations are in regular type, provinces are underlined, and
major islands or island groups are in capital letters. A map of the Republic
of the Philippines is provided with major islands, provinces, and cities noted
(Figure 1).

0L-AD, USAF Occupational and Environmental Health Laboratory, AP0 San
Francisco 96274. Opinions are solely those of the authors and may not reflect
the views of the United States Air Force or the Department of Defense.

2 Current Address: 907 TAG/Aerial Spray Branch, Rickenbacker ANGB,
Columbus, Ohio 43217.

1 .'Brno

REPUBLIC

of the

PHILIPPINES

PROVINCE BOUNDARY
PROVINCE CAPITAL

FIGURE 1

SAMAR

)in
Borongifi

LEYTE

2«3 Ml

Mamoiim

#»

'3fe^

°<s

MINDANAO

DawaoCKy

INDEX TO PROVINCES

1. Bitami

IS. 0«MM

29. Mindin Occitftntil

41. lit ill

54. NbiMs Oriantal

2. Cigtyu

16. ZimbilM

21. Mindin Oriintal

42. Rigrn OccMtiM

55. Aguni dil Rirti

3. Mien Rirti

RTirlac

30. iirinduqui

43. Nigral Ortantal

56. Aguun dil Sir

4. Uveas Syr

11. Bltill

11. Albay

44. Ctbu

57. Linn dil Nirtt

5. Akra

19. Pimpingi

32 Simgin

48 Liyti

SO Liaii dil Sur

6. Riling i-Apiyn

20. Bulicin

33. Rimblii

40. Soutfiam Layta

50. Oukidnin

7. Mountiin

h.ri/ii

34 lubiti

47. Bihil

00. Citibitt

1. Li Uniin

22. Ci»lti

35. Rwfliim timtr

40. Clirtguki

01 Siutti CiUbiti

1 Binguit

23. Ligum

30. Sinwr

49 Surlgti dil lirti

02. Dtvit dil Rirti

10. Itugn

24. Bitingn

37. Eiitim Sunn

50. Surlgit dil Sur

03 Okii dil Sur

U.lnbila

2S. Cimirlnii Rirtt

30. Aklin

51. Zimbungi dil Rirti

04 Divio Oriental

12. Ruivi Vliciyi

20. Cimirinit Sur

39. Cipiz

52. Zimbungi dil Sur

05 Sulu

13. PingiitriM

27. Citinduirtii

40. Antigui

53. Misirnit Occidintil

00. Piliai*

14. Rum Ecija

Mosquito Systematics Vol. 17(1) 1985

SUBFAMILY ANOPHELINAE

Genus Anopheles Meigen

1 . aoaoi Baisas 1946

Clark Air Base, Pampanga , Llavac, Infanta, Quezon , Mt. Banahaw, Los
Banos, Laguna , LUZON.

2. baezai Gater 1933

Los Banos, Laguna, LUZON; Iwahig, PALAWAN; MINDANAO.

3. balevensis Mendoza 1947
Baler, Quezon , LUZON.

4. bengalensis Puri 1930
LUZON.

5. ejercitoi Mendoza 1947
Baler, Quezon , LUZON.

6. fvagilis (Theobald) 1903
LUZON; MINDANAO.

.7. fvanciscoi Reid 1962
Widespread.

8. gigas var. fovmosus Ludlow 1909

Camp John Hay, Benguet , Mountain , LUZON.

9. lestevi Baisas and Ju 1936
Widespread.

10. lindesayi ssp. benguetensis King 1931
LUZON; Mt. Apo, Davao , MINDANAO.

11. manalangi Mendoza 1940
LUZON; MINDANAO.

12. peditaeniatus (Leicester) 1908
Widespread.

13. pilinotwn Harrison and Scanlon 1974
LUZON; MINDANAO; NEGR0S; PALAWAN.

14. pseudobarbirostris Ludlow 1902
Widespread.

15. pseudosinensis Baisas 1935 UTUnnD .
Nueva Ecija , Laguna , LUZON; Naujan, Oriental Mindora , MIND0RA,
Lanao del Sur, MINDANAO.

16. samarensis Rozeboom 1951
SAMAR.

17. vanus Walker 1859

Subic Bay, Zambales , LUZON; Lake Lanao, Lanao del Sur, MINDANAO

Subgenus Cellia Theobald

18. annularis Van Der Wulp 1884
Widespread.

19. balabaoensis balabaoensis Baisas 1936
Balabac Island, PALAWAN.

20. balabaoensis baisasi Col less 1957
Sierra Madre Mountain, LUZON.

21. oristatus King and Baisas 1936

Titunod, Kolambugan, Lanao del Norte , MINDANAO.

22. filipinae Manalang 1930

Spotty deistribution throughout the archipelago.

23. flavirostris (Ludlow) 1914
Widespread.

24. indefinitus (Ludlow) 1904
Widespread.

25. karwari (James) 1902
Widespread.

26. kochi Doenitz 1901
Widespread.

27. kolambuganensis Baisas 1932
Kolambugan, Lanao del Norte , MINDANAO.

28. limosus King 1932
Widespread.

29. litoralis King 1932
Widespread along seashore.

30. ludlowae (Theobald) 1903
Widespread.

31. maaulatus Theobald 1901
Widespread.

Mosquito Systematics Vol. 17(1) 1985

32. mangy anus (Banks) 1906
LUZON; MIND0R0; MINDANAO.

33. parangensis (Ludlow) 1914
Parang, Cotabato , MINDANAO.

34. philippinensis Ludlow 1902
Widespread.

35. riparis King and Baisas 1936
Kolambugan, Lanao del Norte , MINDANAO

36. subpictus Grassi 1899
Widespread.

37. tessellatus Theobald 1901
Widespread.

38. vagus Doenitz 1902
Widespread.

SUBFAMILY CULICINDAE

Genus Aedeomyia Theobald

Important reference: Tyson 1970a.

Subgenus Aedeomyia Theobald

39. catastiota Knab 1909
Widespread.

Genus Aedes Meigen

Important references: Abercrombie 1977. Huang 1977, 1978a, 1978b, 1979.
Knight 1968. Knight and Hull 1951, 1952. Mattingly 1958a, 1958b, 1959, 1961,
1965. Reinert 1970, 1973, 1974, 1976a, 1976b, 1981, 1982. Tyson 1970b.

Subgenus Aedimorphus Theobald

40. alboscutellatus (Theobald) 1905

Northern LUZON; SAMAR; LEYTE; PALAWAN; MINDANAO; MIND0R0; Tawi-Tawi
Island, SULU.

41. oaeeus (Theobald) 1901
Panitian, MIND0R0.

42. lowisii (Theobald) 1910

Calapan, Oriental Mindoro , MIND0R0.

43. pampangensis (Ludlow) 1905

Widespread in LUZON; Mahaplag, LEYTE; San Jose, Occidental Mindoro , MINDORO;
Zamboanga, MINDANAO.

44. punctifemovis (Ludlow) 1921
Central LUZON.

45. vexans (Meigen) 1830
Widespread.

Subgenus Belkinius Reinert

46. aurotaeniatus Edwards 1922

Victoria, Oriental Mindoro , MINDORO; Mt. Siya-Siya, Canlaon Volcano,
Negros Occidental , NEGROS.

Subgenus Canoraedes Edwards

47. ourtipes Edwards 1915

Angeles, Pampanga , LUZON; Iwahig Penal Colony, PALAWAN.

48. masculinus Mattingly 1958
PALAWAN.

49. palawanicus Mattingly 1958
Puerto Princesa, PALAWAN.

Subgenus Ckristophersiomyia Barraud

50. ibis Barraud 1931

San Jose, Nueva Ecija , Alcola, Cagayan , LUZON.

Subgenus Diceromyia Theobald

51. mevonephada (Dyar and Shannon) 1925

LUZON; Osmefia, SAMAR; Tacloban, LEYTE; NEGROS.

Mosquito Systematics Vol. 17(1) 1985

Subgenus Edwavdsaed.es Bel kin

52. impvimens (Walker) 1860

San Jose, Occidental Mindoro , MIND0R0; Puerto Princesa, PALAWAN;
Port Banga, Ft. Pikit, MINDANAO.

Subgenus Finlaya Theobald

53. ananae Knight and Laffoon 1946

Bicol Area, LUZON; Ducong, Basey, Pintanahon, Osmena, SAMAR;
Lago-Lago, Mt. Tobi , Dagami , Tacloban, LEYTE.

54. banksi Edwards 1922 *

Montalban, Rizal , LUZON; Baco , Occidental Mindoro , MINDORO.

55. ovooeus Knight and Laffoon 1946
Subic Bay, Zambales , LUZON.

56. flavipennis (Giles) 1904

Clark Air Base, Pampanga , Mt. Makiling, Los Banos, Laguna , LUZON;
Abuyog, Lago-Lago, Tacloban, LEYTE; Basey River, SAMAR; Pasonanca,
Isabela, Basilan Island, Zamboanga del Sur , MINDANAO.

57. harperi Knight 1948
Subic Bay, Zambales , LUZON

58. jugraensis (Leicester) 1908
Balabac Island, PALAWAN.

59. lacteus Knight 1946

Cape Melville, Balabac Island, PALAWAN.

60. laoagensis Knight 1946
Laoag, Ilocos Norte , LUZON.

61. leucomeves (Giles) 1904

Clark Air Base, Pampanga , LUZON.

62. leuoopleuvus Rozeboom 1946
Irahuan River, PALAWAN.

63. luteus (Ludlow) 1905

Clark Air Base, Pampanga , LUZON

64. luzonensis Rozeboom 1946
Subic Bay, Zambales , LUZON.

65. medlevi Knight and Laffoon 1946 _
Los Banos, Laguna, Jinamoc Island, LUZON; Pintanahon, Osmena,
Calicoan Island, SAMAR; Tacloban, LEYTE; Zamboanga , MINDANAO.

66. melano-ptevus (Giles) 1904 ^

Clark Air Base, Pampanga , Los Barios , Laguna, LUZON; Puerto
Princesa, PALAWAN.

67. niveus (Ludlow) 1903

Subic Bay, Zambales , Batangas , LUZON; Oras, SAMAR; Puerto
Princesa, Irahuan River, Tagburos, PALAWAN.

68. paradissimilis Rozeboom 1946

Cape Melville, Balabac Island, Irahuan River, PALAWAN; Ngolos,
Calicoan Island, SAMAR; Lago-Lago, Baybay, Libyo, Dinagat, Diit
River, Tacloban, LEYTE.

69. ipoioilius (Theobald) 1903
Widespread.

70. vizali (Banks) 1906

Llavac, Infanta, Tayabas, Quezon , LUZON; Canlaon Volcano,
Mt. Siya-Siya, NEGR0S; SAMAR; LEYTE; Lanao , MINDANAO.

71. sapevoi Knight 1946

Subic Bay, Zambales , LUZON.

72. saxiaola Edwards 1922
PALAWAN.

73. sherki Knight 1947 (1948)

Baguio, Benguet, [Knight and Stone 1977 list DavapJ; Baco*,
Occidental Mindoro , MIND0R0.

74. sovsogonensis Banez and Jueco 1966
Sorsogon , LUZON.

75. stonei Knight and Laffoon 1946

Ducong, Basey River, Mt. Lobi , Dagani , LEYTE.

Subgenus Geoskusea Edwards

76. baisasi Knight and Hull 1951

Mangarin Bay, San Jose, Oriental Mindoro , MINDORO; Osmena and
Pintanahon, SAMAR; Iwahig, Penal Colony, PALAWAN; Zamboanga,
MINDANAO.

Subgenus Lorrainea Bel kin

77. amesii (Ludlow) 1903

Pampanga , Los Banos, Laguna , Sorsogon , LUZON; Oras, Calami an
Island, PALAWAN; SAMAR, LEYTE; MINDANAO.

Mosquito Systema tics Vol. 17(1) 1985

78. fwnidus Edwards 1928

Pampanga , LUZON; Culion Island, PALAWAN.

Subgenus Muoidus Theobald

79. fevinus Knight 1947

Subic Bay, Zambales , LUZON; LEYTE; San Ramon Penal Colony Farm,
Zamboanga del Sur , MINDANAO.

80. lanigev (Wiedemann) 1820

Calapan, Oriental Mindoro , MIND0R0; Ludlow Barracks, Petti t
Barracks, Parang, Cotabato , Zamboanga , MINDANAO.

81. quadripunctis (Ludlow) 1910

Dulag, Tacloban, LEYTE; Parang, Cotabato , MINDANAO.

Subgenus Neomelaniconion Newstead

82. Hneatopennis (Ludlow) 1905 * *

Camp Gregg, Bayambang, Pangasinan , Pampanga , Zambales , LUZON.

Subgenus Ochlerotatus Lynch Arriba! zaga

83. vigilax ludlowae (Blanchard) 1905

LUZON; Mangarin, Occidental Mindoro , MINDORO; PANAY; MINDANAO.

Subgenus Paraedes Edwards

84. ostentatio (Leicester) 1908

Mt. Makiling, Laguna and Quezon , LUZON; San Jose, Occidental
Mindoro , MINDORO; Bacungan, PALAWAN.

85. pagei (Ludlow) 1911

San Jose, Occidental Mindoro , MINDORO; Fort Pi kit, Malabang,
Lanao del Sur , San Francisco, Agusan del Sur , MINDANAO.

Subgenus Rhinoskusea Edwards

86. longirostris (Leicester) 1908

Clark Air Base, Pampanga, and Los Banos, Laguna, LUZON; Lapit-Lapit,
Sanga-Sanga, Tawi-Tawi, SULU.

87. wardi Reinert 1976
Widespread.

10

Subgenus Stegomyia Theobald

88. aegypti (Linnaeus) 1762
Widespread.

89. aegypti var. queens landensis (Theobald) 1901

Balian , Pangil , College of Agriculture*, Laguna , LUZON; Bongao*
Island, SULU.

90. albolineatus (Theobald) 1904

Pampanga , Los Banos, Laguna , LUZON; SAMAR; Balabac Island, PALAWAN,

91. albopictus (Skuse) 1894
Widespread.

92. aloasidi Huang 1972
Widespread.

93. arborioola Knight and Rozeboom 1946
Shohoton Springs, Basey River, SAMAR.

94. bconbusicola Knight and Rozeboom 1946
Culion Island, PALAWAN.

95. boharti Knight and Rozeboom 1946

Laguna*, LUZON; Baco*, Occidental Mindoro , MIND0R0; Osmena, SAMAR.

96. desmotes (Giles) 1904

Clark Air Base, Pampanga , LUZON; LEYTE; NEGR0S; PALAWAN.

97. gardnerii (Ludlow) 1905
Widespread.

98. hoogstraali Knight and Rozeboom 1946
Subic Bay, Zambales , LUZON.

99. impatibilis (Walker) 1859

San Jose, Occidental Mindoro , MINDORO.

100. laffooni Knight and Rozeboom 1946

San Ramon, Zamboanga del Sur , MINDANAO.

101. mediopunctatus (Theobald) 1905
Zambales , LUZON; PALAWAN.

102. paullusi Stone and Farner 1945
Widespread.

103. platylepidus Knight and Hull 1951

Puerto Princesa, Cape Melville, Balabac Island, PALAWAN.

Mosquito Systematics Vol. 17(1) 1985 11

Subgenus Vevvallina Theobald

104. adustus Laffoon 1946

Cape Melville, Balabac Island, PALAWAN.

105. butleri Theobald 1901

Tacloban and Palo, LEYTE; Culion Island, Iwahig, Basbas Island,
Puerto Princesa, PALAWAN.

106. comply lo stylus Laffoon 1946

Quezon , LUZON; San Jose, Occidental Mindoro , MIND0R0; Tacloban, LEYTE:
Balsahan, Irahuan River, Cape Melville, Balabac Island, PALAWAN; San
Ramon, Zamboanga del Sur , MINDANAO.

107. dux Dyar and Shannon 1925

Clark Air Base, Pampanga , Subic Bay, Zambales , Camp Nichols, Rizal ,
San Jose, Bulacan , Manila, LUZON; Caminawit Point, MINDORO; SAMAR;
Zamboanga , MINDANAO.

108. hamistylus Laffoon 1946

San Jose, Occidental Mindoro , MINDORO; Cape Melville, Balabac
Island, PALAWAN.

109. Qohnsoni Laffoon 1946

Tanaun, LEYTE; Isabela, Basilan Island, Pasonanca, San Ramon,
Zamboanga del Sur , MINDANAO.

110. maorodixoa Dyar and Shannon 1925

Tayabas, Infanta, Quezon , LUZON; San Jose, Occidental Mindoro ,
MINDORO; Tacloban, LEYTE; Kabakan Mercedes, Parang, Cotabato ,
MINDANAO.

111. margarsen Dyar and Shannon 1925

Subic Bay, Zambales , Camp Eldridge, Los Banos, Laguna , Tungking,
Mangga, San Jose, Bulacan , LUZON; San Jose, Kabakan, Zamboanga
del Norte , MINDANAO.

112. nigrotarsis (Ludlow) 1908

Subic Bay, Zambales , Infanta, Quezon , Camp Nicols, Wack-Wack,
Rizal , Calauan, Laguna , LUZON; Osmena, SAMAR; Abuyog, Tacloban,
LEYTE; San Ramon, Zamboanga del Sur , MINDANAO.

113. nubioolus Laffoon 1946

Sibulan River, Mt. Apo, Davao , MINDANAO.

114. panayensis Ludlow 1914

Iloilo, PANAY; Calicoan Island, SAMAR; Culion, San Pedro, Cape
Melville, Balabac Island, PALAWAN; Basbas, Zamboanga , MINDANAO.

12

115. philippinensis Delfinado 1968
Mahaplag, LEYTE.

116. robertsi Laffoon 1946
Tacloban, LEYTE.

117. uncus (Theobald) 1901

Irahuan River, Puerto Princesa, Cape Melville, Balabac Island,
Culion Island, PALAWAN; Cotabato, MINDANAO.

Genus Avmigeres Theobald
Important references: Baisas 1965. Delfinado 1966. Thurman 1959

Subgenus Armigeres Theobald

118. apoensis Bohart and Farner 1944
Sibulan River, Mt. Apo, Davao , MINDANAO.

119. aureolineatus (Leicester) 1908
Mantalingajan, Pinigisan, PALAWAN.

120. azurini Basio 1971

Mt. Makiling, Laguna , LUZON.

121. baisasi Stone and Thurman 1958
Widespread.

122. ejevoitoi Baisas 1935

Baguio, Benguet , LUZON; Minaluang and Bal-Ason, Misamis Oriental ,
Mt. McKinley, Davao , Daragawan, Cotabato , San Francisco, Agusan ,
MINDANAO.

123. joloensis (Ludlow) 1904

LUZON; LEYTE; SAMAR; MIND0R0; PALAWAN; MINDANAO; SULU

124. malayi (Theobald) 1901

Bulacan , Laguna , and Rizal , LUZON; NEGR0S ORIENTAL; LEYTE
Calicoan Island, SAMAR; PALAWAN; Misamis Oriental , Davao ,
Cotabato , MINDANAO; Siasi Island, SULU.

125. manalangi Baisas 1935

Bulacan , Laguna , LUZON; San Jose, Occidental Mindoro , MIND0R0;
Mt. Empagatao, MINDANAO.

126. setifev Delfinado 1966
Mantalingajan, Pinigisan, PALAWAN.

Mosquito Systematics Vol. 17(1) 1985 13

127. subalbatus (Coquillett) 1898

LUZON; SAMAR; PALAWAN; MINDANAO; JOLO.

Subgenus Leioesteria Theobald

128. digitatus (Edwards) 1914

Laguna , LUZON; San Jose, Occidental Mindoro , MINDORO; Bal-Ason,
NEGROS ORIENTAL; Lanao , MINDANAO.

129. flavus (Leicester) 1908

Mt. Banahaw, Lucban, Tayabas, Quezon , LUZON.

130. magnus (Theobald) 1908

La Union , Batangas , LUZON; San Jose, Occidental Mindoro ,
MINDORO; MINDANAO.

131. omissus (Edwards) 1914

San Jose, Occidental Mindoro , MINDORO.

Genus Coquillettidia Dyar
Important reference: Delfinado 1966.

Subgenus Coquillettidia Dyar

132. aureosquammata (Ludlow) 1909

Parang, Ludlow Barracks, Cotabato , MINDANAO.

133. crassipes (Van der Wulp) 1881

Central LUZON; San Jose, Occidental Mindoro , MINDORO; Lake Maim tin,
Surigao , Ludlow Barracks, Parang, Fort Pi kit, Cotabato , Petti t
Barracks, Zamboanga , MINDANAO.

134. nigrosignata (Edwards) 1917

Siasi Island, Tawi-Tawi Island, Tarakan, North Batu-Batu, SULU.

135. ochraoea (Theobald) 1903

Clark Air Base, Pampanga ; Los Banos, Laguna , LUZON; San Antonio,
SAMAR; Tacloban, LEYTE; Ludlow Barracks, Cotabato , MINDANAO.

Genus Culex Linnaeus

Important references: Cagampang-Ramos 1979. Delfinado 1966.
Sirivanakarn 1972, 1975, 1976, 1977a, 1977b.

14

Subgenus Culex Linnaeus

136. alls Theobald 1903

Subic Naval Base, Zambales , LUZON; Osmena, SAMAR.

137. annulivostvis Skuse 1889
Widespread.

138. bitaeniorhynchus Giles 1901
Widespread.

139. edwardsi Barraud 1923

John Hay Air Base, Baguio, Benguet , Clark Air Base, Pampanga,
LUZON.

140. fasyi Baisas 1938

Benguet Otok, Chico River, Bontok, Mountain , Baguio City,
Benguet, LUZON.

141. fusooaephala Theob al d 1907
Widespread.

142. gelidus Theobald 1901
Widespread.

143. inoognitus Baisas 1938
Widespread.

144. infula Theobald 1901

Central LUZON; Cadiz, NEGR0S; Rizal, Palo, Lago-Lago, Baybay,
Dulag, Loreto, Dinagut, Tacloban, Pinamuan Gulf, Jinamoc Island,
LEYTE: MINDANAO; Tawi-Tawi Island, SULU.

145. luzonensis Sirivanakarn 1976

Dal ton, Imugan, Nueva Vizcaya , Baguio, Benguet , Mountain ,
Subic Naval Base, Zambales , LUZON.

146. mimulus Edwards 1915

Tacloban, LEYTE; Balabac, Puerto Princesa, Tagburos, Iwahig,
Irahuan River, Balsahan River, Bacungan, PALAWAN; San Francisco,
Agusan , Isabela, Basilan Island, Zamboanga del Sur , MINDANAO;
Muni, Jolo Island, SULU.

147. ovientalis Edwards 1921

Laguna , LUZON; Balabac Island, PALAWAN.

148. philippinensis Sirivanakarn 1976

Tay-Tay, Rizal , LUZON; Osmena, SAMAR; Zamboanga , MINDANAO.

149. pseudovishnui Col less 1957
Widespread.

Mosquito Systematics Vol. 17(1) 1985 15

150. quinquefasoiatus Say 1823
Widespread.

151. sinensis Theobald 1903

Wack-Wack, Rizal , Pampanga , Bulacan, LUZON; Calacad, SAMAR.

152. sitiens Wiedemann 1828
Widespread.

153. tritaeniorhynohus Giles 1901
Widespread.

154. vishnui Theobald 1901
Widespread.

155. whitei Barraud 1923

San Fabian, Pangasinan , Subic Naval Base, Zambales , Laguna ,
Clark Air Base*, Pampanga , LUZON; Mt. Lobi , LEYTE.

156. whitmorei (Giles) 1904

Baguio, Benguet , Zambales , Pampanga , Rizal , Laguna , LUZON;
San Jose, Occidental Mindoro , MIND0R0; Guirang and Calacad,
SAMAR; LEYTE.

Subgenus Culioiomyia Theobald

157. delfinadoae Sirivanakarn 1973
Tawi-Tawi, Sangir Island, SULU.

158. fragilis Ludlow 1903

Camansi, Santo! , La Union, Subic Bay, Zambales , LUZON; San jose,
Occidental Mindoro , MINDORO; SAMAR; Carigara, Jinamoc Island,
LEYTE; Puerto Princesa, PALAWAN; San Francisco, Agusan , Parang,
Cotabato , MINDANAO.

159. nigvopunctatus Edwards 1926

La Union , Pampanga , and Zambales , LUZON: Jinamoc Island and
Leyte Gulf, LEYTE; Osmena, SAMAR; Bacungan and Iwahig Penal
Colony, PALAWAN; Parang, Cotabato , and San Ramon, Zamboanga
del Sur , MINDANAO.

160. pallidothorax Theobald 1905

San Jose, Occidental Mindoro , MINDORO.

161. papuensis (Taylor) 1914

Puerto Princesa, PALAWAN; Mt. McKinley, Davao , MINDANAO.

16

162. spathifuroa (Edwards) 1915

Caminawit Pt. and Calapan, Oriental Mindoro , MINDORO; Abuyog and
Jinamoc Island, LEYTE; Calicoan Island, SAMAR; Parang, Cotabato ,
MINDANAO.

Subgenus Eumelanomyia Theobald

163. baisasi Sirivanakarn 1972

Osmena, SAMAR; Kolambugan, Lanao del Norte , MINDANAO.

164. brevipalpis (Giles) 1902

Agoo, Balaoan, Camansi , La Union , Baguio City, Benguet , Subic
Bay, Zambales , Mt. Makiling, Laguna , LUZON; San Jose, Occidental
Mindoro , MINDORO; Irahuan River, Rosa, PALAWAN; MINDANAO.

165. oataractarum Edwards 1923
Mt. Makiling, Laguna , LUZON.

166. foliatus Brug 1932

Subic Naval Base, Zambales , Dingalan Bay, Tayabas, Quezon , LUZON;
San Jose, Occidental Mindoro , MINDORO; Osmena, SAMAR; Baybay,
Lago-Lago, LEYTE; Iwahig, Puerto Princesa, PALAWAN; Parang,
Cotabato , Kolambugan, Lanao del Norte , MINDANAO.

167. hinglungensis Chu 1957

San Jose, Occidental Mindoro , MINDORO; San Pedro, Puerto Princesa,
Culion Island, PALAWAN.

168. lati foliatus Delfinado 1966

Mt. Makiling, Laguna , LUZON; San Jose, Occidental Mindoro , MINDORO;
Balsahan River, Cabayugan, Marble Mountain, Panitan, PALAWAN.

169. lauveli Baisas 1935

San Jose, Nueva Ecija , Subic Naval Base, Zambales , Batangas , San
Pablo City, Calauan, Laguna , LUZON; San Jose, Occidental Mindoro ,
MINDORO; San Antonio, Osmena, Guirang, Sohotan, SAMAR; Baybay,
Lago-Lago, Tacloban, LEYTE; Iwahig, Puerto Princesa, PALAWAN;
Mercedes, Quimpot, Zamboanga , Bukidnon , Lanao , MINDANAO.

170. okinawae Bohart 1953

Trinidad, Benguet , San Fernando, La Union , Mountain ,
LUZON.

171. unoinatus Delfinado 1966
Osmena, SAMAR.

172. yeageri Baisas 1935

Iwahig, Bacungan, Quezon, Panitan, PALAWAN.

Mosquito Systematics Vol. 17(1) 1985 17

Subgenus Lophoaevaomyia Theobald

173. oinotellus Edwards 1922

LUZON; San Antonio, SAMAR; Palo, Dulag, Diit River, Buruguin, LEYTE.

174. oubitatus Coll ess 1965

Puerto Princesa, PALAWAN; San Francisco, Agusan , MINDANAO.

175. fullevi (Ludlow) 1909

Ludlow Barracks, Parang, Kabakan, Cotabato , San Francisco, Agusan ,
MINDANAO; Jolo Island, SULU.

176. gibbulus Delfinado 1966

San Jose, Occidental Mindoro, MIND0R0; Brooks' Pt. , Uring-Uring,

PALAWAN.

177. infantulus Edwards 1922

Agoo, La Union, Subic Naval Base, Zambales , Mt. Makiling, Laguna,
LUZON; San Jose, Occidental Mindoro , MINDORO; Osmena, SAMAR; Palo,
Lago-Lago, Baybay, LEYTE; Parang, Cotabato , MINDANAO.

178. josephineae Baisas 1935

San Jose, Occidental Mindoro , MINDORO; Osmena, SAMAR; Santa Rosa,
Tacloban, Diit, Lago-Lago, Baybay, Dagamit, and Mt. Lobi , LEYTI;
Ludlow Barracks, Parang, and Kabakan, Cotabato , Pasonanca, Zamboanga
del Sur , San Francisco, Agusan, San Francisco, Surigao , MINDANAO.

179. kuhnsi King and Hoogstraal 1955

Baguio City, Benguet , Dal ton, Nueva Vizcaya , Subic Bay, Zambales ,
Mt. Makiling, Laguna , Calaca, Batangas , LUZON; San Jose, Occidental
Mindoro , MINDORO; Fabrica, Negros Occidental , Balinsayao, Lago-Lago
Bay, LEYTE; Kolambugan, Lanao del Norte , Mt. Apo, Davao , Wenes,
Cotabato , MINDANAO; Mampallan, Talipaw, Indanan, JOLO.

180. lavatae Stone and Bohart 1944

Baguio City, Benguet , Dalton, Nueva Vizcaya , Munoz, San Jose,
Nueva Ecija , Subic Naval Base, Zambales , Mt. Makiling, Laguna ,
LUZON; San Jose, Occidental Mindoro , MINDORO; Osmena, SAMAR;
Tacloban, LEYTE; Mt. Malino, Irahuan River, Puerto Princesa,
PALAWAN; Indanan, JOLO.

181. maodonaldi Colless 1965

San Jose, Occidental Mindoro , MINDORO.

182. mammilifer (Leicester) 1908

Subic Naval Base, Zambales , Mt. Makiling, Laguna , LUZON; San Jose,
Occidental Mindoro , MINDORO; Osmena, SAMAR; Dagami , Lago-Lago Bay,
IFYTE: Ginqoog, Misamis Oriental , Kolambugan, Lanao del Norte,
Davao, MINDANAO; Iwahig, Irakman, Balsahan River, PALAWAN.

18

183. minor (Leicester) 1908

Trinidad, Mountain , Dal ton, Nueva Vizcaya , Tayabas, Quezon ,
Subic Naval Base, Zambales , Mt. Isarog, Camarines Sur , LUZON;
Osmefia, Calaccad, Sohoton, SAMAR; Lago-Lago, Baybay, LEYTE;
Mat Mali no, Irahuan River, Iwahig, Puerto Princesa, PALAWAN;
Jolo Island, SULU.

184. paraeuleatus Sirivanakarn 1977

Diit River, Tacloban, LEYTE; Parang, Cotabato , MINDANAO.

185. quadripalpis (Edwards) 1914

Mt. Makiling, Laguna , LUZON; Puerto Princesa, Iwahig, Cabayuyan,
Ludlow Barracks, PALAWAN.

186. reidi Col less 1965
Uring-Uring, Brooks Point, PALAWAN.

187. rubithoraois (Leicester) 1908

Subic Naval Base, Zambales , LUZON; San Jose, Occidental Mindoro ;
MIND0R0; Tacloban, Palo, LEYTE; Jolo Island, SULtr

Subgenus Lutzia Theobald

188. fusoanus Wiedemann 1820
Widespread.

189. halifaxii Theobald 1903

Subic Bay, Zambales , Clark Air Base, Pampanga , Calauan, Laguna ,
LUZON; San Jose, Occidental Mindoro , MINDORO; Sohoton Spring,
Osmena, SAMAR; Lake Mai nit, Surigao , San Francisco, Agusan ,
Pasonanca, San Ramon, Zamboanga del Sur, Cotabato , MINDANAO;
Cape Melville, Balabac Island, Culion Island, Irahuan River,
PALAWAN.

Genus Fioalbia Theobald

Important reference: Mattingly 1957,

190. ludlowaeBrunetti 1920
Cudarangan, MINDANAO.

Genus Heizmannia Ludlow
Important reference: Mattingly 1970.

Mosquito Systematics Vol. 17(1) 1985 19

Subgenus Eeizmannia Ludlow

191. scintillans Ludlow 1905

Clark Air Base, Pampanga , LUZON.

Genus Eodgesia Theobald

Important reference: Delfinado 1966.

192. malayi Leicester 1908

Los Banos, Laguna , LUZON; San Jose, Occidental Mindoro , MINDORO;
Santa Rosa, LEYTE; Titunod Creek, Kolambugan, Lanao del Norte ,
MINDANAO.

193. quasi sanguinae Leicester 1908

San Jose, Occidental Mindoro , MINDORO; Lanao, San Francisco,
Agusan , MINDANAO.

Genus Malaya Leicester

Important reference: Wharton 1947.

194. genurostris Leicester 1908

Clark Air Base, Pampanga , LUZON; Ludlow Barracks, Parang
Cotabato , MINDANAO.

195. leei (Wharton) 1947

Ludlow Barracks, Cotabato , MINDANAO.

Genus Mansonia Blanchard
Important references: Delfinado 1966. Wharton 1978.

Subgenus Mansonioides Theobald

196. annulata Leicester 1908

Ludlow Barracks, Ft. Pikit, Cotabato , MINDANAO.

197. annulifeva (Theobald) 1901

Clark Air Base, Pampanga , San Fabian, Mangit, Batangas , Camp
Eldridge, Los Banos, Laguna , Ft. McKinley, Rizal , Mountain ,
LUZON; Reina Regente, Ludlow Barracks, Parang, Cudarangan,
Cotabato, MINDANAO.

20

198. bonneae Edwards 1930

Clark Air Base, Pampanga , LUZON; Gungnam, Marirong, Kambing,
PALAWAN.

199. dives (Schiner) 1868

San Jose, Occidental Mindoro , MIND0R0; San Francisco, Agusan,
MINDANAO.

200. uniformis (Theobald) 1901
Widespread.

Genus Mimomyia Theobald
Important references: Delfinado 1966. Mattingly 1971. White 1974

Subgenus Etovleptiomyia Theobald

201. elegans (Taylor) 1914
Kabakan, Cotabato , MINDANAO.

202. luzonensis (Ludlow) 1905

Bayambang, Pangasinan , Subic Naval Base, Zambales , Camp Nichols,
Rizal, Pampanga , LUZON; San Jose, Occidental Mindoro , MINDORO;
Kabakan, Cotabato , MINDANAO.

Subgenus Ingramia Edwards

203. deguzmanae (Mattingly) 1957

Baguio City, Benguet , Cagayan , LUZON; San Jose, Occidental
Mindoro , MINDORO; NEGR0S; Mt. McKinley, Davao , Lanao, Surigao,
MINDANAO. *~

Subgenus Mimomyia Theobald

204. ohamberlaini Ludlow 1904

Munoz Agricultural School, Nueva Ecija , Natividad, San Miguel,
Tayug, Pangasinan , Rizal , Calauan, Laguna , LUZON; San Jose,
Occidental Mindoro , MINDORO; LEYTE.

205. ohamberlaini issp. metallioa (Leicester) 1908
LUZON.

206. hybrida (Leicester) 1908

Munoz Agricultural School, Nueva Ec ija, LUZON; Cudarangan,
MINDANAO.

Mosquito Systematics Vol. 17(1) 1985 21

Genus Orthopodomyia Theobald

Important reference: Zavortink 1971.

207. andamanensis Barraud 1934

Cape Melville, Balabac Island, PALAWAN.

208. anopheloides (Giles) 1903

Bagnio, Benguet , Mountain , Laguna , LUZON; San Jose,
Occidental Mindoro , MIND0R0; NEGROS; LEYTE; PALAWAN.

209. madvensls Baisas 1946

Mt. Isarog, Camarines Sur , Baguio, Benguet , Laguna , LUZON; PALAWAN.

Genus Topomyia Leicester
Important references: Delfinado 1966. Miyagi 1980.

Subgenus Suaymyia Thurman

210. apsarae Klein 1977

Montible, Puerto Princesa, PALAWAN.

211. imitata Baisas 1946
Kidapawan, Cotabato , MINDANAO.

Subgenus Topomyia Leicester

212. avgyropalpis Leicester 1908

Ludlow Barracks, Parang, Cotabato , Davao , MINDANAO.

213. barbus Baisas 1946
MINDANAO.

214. oabvevai Miyagi, Toma, and Rivera 1983
Mt. Apo, Davao , MINDANAO.

215. dejesusi Baisas and Feliciano 1953
Mt. Apo, Davao , MINDANAO.

216. hernandoi Baisas and Feliciano 1953
Mt. Apo, Davao , MINDANAO.

217. pseudobarbus Baisas 1946

San Miguel, Bulacan , Subic Bay, Zambales , LUZON; Puerto Princesa,

PALAWAN.

22

Genus Triptevoides Giles

Important references: Baisas and Ubaldo-Pagayon 1949, 1952
Mattingly 1980, 1981.

Subgenus Rachionotomyia Theobald

218. vozebooml Baisas and Ubaldo-Pagayon 1953
Pasonanca, Zamboanga del Sur , MINDANAO.

Subgenus Tvicholeptomyia Dyar and Shannon

219. apoensis Baisas and Ubaldo-Pagayon 1953

Sumilao, Maluko, Tagalaon, Bukidnon , Taglawig, Tagum, Mt. McKinley,
Mt. Apo, Davao , MINDANAO.

220. bavvaudi Baisas and Ubaldo-Pagayon 1953

Famy, Laguna , Sierra Madre Mountain, Baler, Quezon, Mountain,
LUZON.

221. belkini Baisas and Ubaldo-Pagayon 1953
Bacungan, Puerto Princesa, PALAWAN.

222. ohristophersi Baisas and Ubaldo-Pagayon 1953,
Baler, Quezon , Mountain , LUZON.

223. delpilari Baisas and Ubaldo-Pagayon 1953

Mt. Makiling, Los Banos, Laguna , Llavac, Infanta, Baler, Quezon,
LUZON.

224. miorooala (Dyar) 1929

Bamban River, Clark Air Base, Pampanga , along Banaue-Bontoc Road,
Mountain , Llavac, Infanta, Baler, Quezon , LUZON.

225. nepenthicola (Banks) 1909

Trindad and along Banaue-Bontoc Road, Mountain , Llavac, Infanta,
Baler, Quezon , Mt. Makiling, Los Banos, Famy, Laguna , LUZON.

226. woxasi Baisas and Ubaldo-Pagayon 1953
Sumilao, Bukidnon , MINDANAO.

227. werneri Baisas and Ubaldo-Pagayon 1953
Mt, Apo, Mt. McKinley, Davao , MINDANAO.

Subgenus Tripteroides Giles

228. antennalis Bohart and Farner 1944

Sibulan River, Mt. Apo, Mt. McKinley, Davao , MINDANAO.

Mosquito Systematics Vol. 17(1) 1985 23

229. claggi Bohart and Farner 1944

Sibulan River, Mt. Apo, Davao , MINDANAO.

230. dyari Bohart and Farner 1944

Laguna , Quezon , LUZON; Davao , MINDANAO.

231. dyi Baisas and Ubaldo-Pagayon 1953
Mt. McKinley, Davao , MINDANAO.

232. evlindae Baisas and Ubaldo-Pagayon 1953
Mt. McKinley, Davao , MINDANAO.

233. hoogstraali Baisas 1947
Mt. Data, Mountain , LUZON.

234. indeterminatus Baisas and Ubaldo-Pagayon 1953
Mt. McKinley, Davao , MINDANAO.

235. intermediatus Baisas and Ubaldo-Pagayon 1953
Taglawig, Tagum, Davao , MINDANAO.

236. knighti Baisas and Ubaldo-Pagayon 1953
Pasonanca, Zamboanga del Sur , MINDANAO.

237. mabinii Baisas and Ubaldo-Pagayon 1953
Cotabato, Lanao , MINDANAO.

238. malvari Baisas and Ubaldo-Pagayon 1953
Laguna , Quezon , LUZON.

239. monetifev (Dyar) 1920
Widespread.

240. nitidoventer (Giles) 1904
Widespread.

241. powelli (Ludlow) 1909

LUZON; SAMAR; LEYTE; Balabac Island, PALAWAN.

242. powelli esoodae Baisas and Ubaldo-Pagayon 1953

Mt. McKinley, Davao , MINDANAO; Cape Melville, Puerto Princesa,
PALAWAN; J0L0.

243. powelli laffooni Baisas and Ubaldo-Pagayon 1953
SAMAR; MINDANAO.

244. powelli mattinglyi Baisas and Ubaldo-Pagayon 1953
Irahuan River, Culion, PALAWAN.

245. veiseni Basio 1971

Mt. Makiling, Laguna , LUZON.

24

246. riverai Miyagi , Toma, and Tsukamoto 1983
Iwahig, PALAWAN.

247. simulatus Baisas and Ubaldo-Pagayon 1953
Banaue, Mountain , LUZON.

248. sullivanae Baisas and Ubaldo-Pagayon 1953
Manatanao, Davao , MINDANAO.

249. toffaletii Baisas and Ubaldo-Pagayon 1953
Iwahig, Culion, PALAWAN.

250. uiohanooi Baisas and Ubaldo-Pagayon 1953
Mt. McKinley, Davao, MINDANAO.

Genus Uranotaenia Lynch Arribalzaga

Important references: Delfinado 1966. Peyton 1972, 1977. Peyton
and Klein 1970. Peyton and Rattanari thikul 1970.

Subgenus Pseudoficalbia Theobald

251. abstrusa Peyton 1977

San Jose, Occidental Mindoro , MIND0R0.

252. bioolor Leicester 1908

Burgos, Mountain , Los Banos, Laguna , Limay, Bataan , LUZON;
Kidapawan, Cotabato , MINDANAO.

253. confusa Peyton 1977

San Jose, Occidental Mindoro , MINDORO.

254. demeilloni Peyton and Rattanari thikul 1970
Baguio, Benguet , LUZON.

255. harrisoni Peyton 1977

San Antonio, Oriental Mindoro , MINDORO.

256. modesta Leicester 1908

Calauan, Laguna, LUZON; Kolambugan, Lanao del Norte , MINDANAO.

257. obsouva Edwards 1915

Burgos, Mountain , LUZON; San Jose, Occidental Mindoro , MINDORO;
K 98 Creek, Ludlow Barracks, Parang, Cotaba to, San Francisco,
Ausan , MINDANAO.

258. pylei Baisas 1946

Llavac, Laguna , LUZON; Kidapawan, Cotabato , MINDANAO.

Mosquito Systematics Vol. 17(1) 1985 25

259. rossi Delfinado 1966

San Jose, Occidental Mindoro , MINDORO.

Subgenus Uranotaenia Lynch Arriba Izaga

260. annandalei Barraud 1926

San Jose, Occidental Mindoro , MINDORO; Titunod Creek, Kolambugan,
Lanao del Norte , Parang, Cotabato , MINDANAO.

261. avguellesi Baisas 1935

Calauan, Laguna , La Union , Clark Air Base*, Pampanga , LUZON;
San Jose, Occidental Mindoro , MINDORO; Tacloban, LEYTE.

262. argyrotavsis Leicester 1908

San Jose, Calicoan Island, Occidental Mindoro , MINDORO; Kabakan,
Parang, Cotabato , MINDANAO; Iwahig, PALAWAN.

263. olava Dyar and Shannon 1925

San Jose, Occidental Mindoro , MINDORO; Parang, Salimbao, Ludlow
Barracks, Cotabato , MINDANAO.

264. faloipes Banks 1906

Clark Air Base, Pampanga , Bulacan , Rizal , Manila, LUZON; San Jose
Occidental Mindoro , MINDORO; Kidapawan, Cotabato , MINDANAO.

265. heiseri Baisas 1935

Parang and Kabakan, Cotabato , MINDANAO.

266. lateralis Ludlow 1905

Camp Eldridge, Laguna , Camp Nichols, Rizal , LUZON; San Jose,
Occidental Mindoro , MINDORO; Tacloban, LEYTE; Kolasi , Kolambugan,
Lanao del Norte , Jinamoc Island, Caminawit Ft., Cotabato , MINDANAO.

267. ludlowae Dyar and Shannon 1925

Laguna , LUZON; Ludlow Barracks, Parang, Simoay, Cotabato ,
MINDANAO.

268. mendiolai Baisas 1935

Olongapo Road, Zambales , Los Banos, Laguna , Atimonan, Quezon ,
Bataan , LUZON; San Jose, Occidental Mindoro , MINDORO; Kabakan
Kolambugan, Lanao del Norte , MINDANAO.

269. metatavsata Edwards 1914
Camp Nichols, Rizal , LUZON.

270. nivea Leicester 1908

San Jose, Occidental Mindoro , MINDORO; Ft. Pikit, Kabakan, Parang,
Ludlow Barracks, Reina Regente, Cotabato , MINDANAO.

271. reyi Baisas 1935

Simoay, Cotabato, MINDANAO.

26

Genus Zeugnomyia Leicester

Important reference: Delfinado 1966.

272. aguilavi Baisas and Feliciano 1953
Mt. Apo, Davao , MINDANAO.

273. fajardoi Baisas and Feliciano 1953
Mt. Apo, Davao , MINDANAO.

274. gracilis Leicester 1908
Ube, Laguna , LUZON.

275. lawtoni Baisas 1946

Llavac, Infanta, Quezon , LUZON; Wenes, Calinan, Davao , MINDANAO.

SUBFAMILY TOXORHYNCHITINAE
Genus Toxovhynohites Theobald
Important reference: Belkin 1962.

Subgenus Toxovhynohites Theobald

276. amboinensis (Doleschall) 1857

MIND0R0; Hacienda Louisiana, Mailum, NEGROS; Guimaras Island,
CEBU; Galong and Iloilo, PANAY; Margosatubig, Zamboanga del Sur,
MINDANAO.

277. gigantulus (Dyar and Shannon) 1925

Clark Air Base, Pampanga , Limay, Bataan , San Andres, Rizal ,
Laguna , LUZON.

278. nepenthis (Dyar and Shannon) 1925
Los Banos, Laguna , LUZON.

279. splendens (Wiedemann) 1819

Clark Air Base, Pampanga , Los Banos, Laguna , San Andres, Rizal,
LUZON; NEGROS; CEBU; PALAWAN.

Mosquito Systematics Vol. 17(1) 1985 27

ACKNOWLEDGMENTS

We wish to thank Mr. R. Sese, Mr. J. Velasquez, Ms. F. Butiu, Mrs. P.
Enriquez, and Ms. M. Carlos, for their assistance and extraordinary patience
during preparation of the manuscript.

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